By Glenn D. Prestwich, Gary J. Blomquist
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Am. 58, 6 0 4 608. Silk, P. , Juenen, L. P. , and Lonergan, G. C. (1985). A biologically active analogue of the primary sex-pheromone components of spruce budworm, Choristoneura fumiferana (Lepidop tera: Noctuidae). Can. Entomol 117, 2 5 7 - 2 6 0 . Silverstein, R. M. (1979). Enantiomeric composition and bioactivity of chiral semiochemicals in insects. In "Chemical Ecology: Odor Communication in A n i m a l s " (F. J. ), pp. 113-146. Elsevier, Amsterdam. Silverstein, R. M . , Rodin, J. O . , and Wood, D .
Percy-Cunninghann a n d J . A . M a c D o n a l d 42 which are found at the bases of the second valvifers and on the ninth tergite. Morphological studies, reinforced by behavioral studies, led Tagawa (1977, 1983) to conclude that the glandular fields associated with the second valvifers were the sites of production and release of the sex pheromone. The glands are composed of two types of glandular cells: modified trichogen cells with associ ated setae and ducted columnar cells. These results contradict those of Weseloh (1980), who stated that glands on the eighth tergite (equivalent to the ninth tergite) actually produced the pheromone.
Abdomen—Genital Sex pheromone production and release is often associated with the genital region of the abdomen. The glands of this region may be associated with glands in another part of the body, as the scent brushes of male Lepidoptera. However, epidermal glands which appear to be the sole source of the pheromones have also been identified in several orders. Scoφion flies of the genus Panorpa have a ventral glandular patch on the genital bulb which projects anterioriy into the bulb but is eversible.